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Echinochloa crus-galli (L.) P. Beauv.

Family :

Famille :

Poaceae

Synonym(s) :

Synonyme(s) :

Panicum crus-galli L., basionym (USDA-ARS 2024)
Echinochloa disticha St.-Lag. (POWO 2024)
Milium crus-galli (L.) Moench (POWO 2024)
Oplismenus crus-galli (L.) Dumort. (POWO 2024)
Orthopogon crus-galli (L.) Spreng. (POWO 2024)
Panicum alectorocnemum St.-Lag. (POWO 2024)
Panicum alectromerum Dulac (POWO 2024)
Panicum crus-galli subsp. breviaristatum Ehrh. (POWO 2024)
Panicum grossum Salisb. (POWO 2024)
Pennisetum crus-galli (L.) Baumg. (POWO 2024)

Common Name(s) :

Nom(s) commun(s) :

Barnyardgrass

(English) (AOSA 2024; USDA-NRCS 2024)
Barnyard grass (English) (Wiersema and Léon 1999; Darbyshire 2003; Rojas-Sandoval and Acevedo-Rodriguez 2014; USDA-ARS 2024)
Barnyard millet (English) (Rojas-Sandoval and Acevedo-Rodriguez 2014; USDA-ARS 2024)
Cocksfoot grass (English) (USDA-ARS 2024)
Cockshin grass (English) (USDA-ARS 2024)
Cockspur (English) (Rojas-Sandoval and Acevedo-Rodriguez 2014; USDA-ARS 2024)
Cockspur grass (English) (Rehm 1994; Rojas-Sandoval and Acevedo-Rodriguez 2014)
Cockspur-panic (English) (USDA-ARS 2024)
Water grass (English) (USDA-ARS 2024)
Deneiba (transcribed Arabic) (Boulos and el-Hadidi 1984)
Echinochloa pied-de-coq (French) (Barkworth et al. 2003; Rojas-Sandoval and Acevedo-Rodriguez 2014)
Panic pied-de-coq (French) (Rojas-Sandoval and Acevedo-Rodriguez 2014; USDA-ARS 2024)
Pied de coq (French) (Rehm 1994; Rojas-Sandoval and Acevedo-Rodriguez 2014)
Échinochloa pied-de-coq (French) (Darbyshire 2003)
Gewöhnliche Hühnerhirse (German) (USDA-ARS 2024)
Hühnerhirse (German) (Rehm 1994)
Sanwak (India) (Rehm 1994)
Giavone (Italian) (Rojas-Sandoval and Acevedo-Rodriguez 2014)
Canarana (Portuguese) (Rojas-Sandoval and Acevedo-Rodriguez 2014)
Capim-andrequicé (Portuguese) (Rojas-Sandoval and Acevedo-Rodriguez 2014)
Miha-pe-de-galo (Portuguese) (Rojas-Sandoval and Acevedo-Rodriguez 2014)
Arrocillo (Spanish) (Rojas-Sandoval and Acevedo-Rodriguez 2014; USDA-ARS 2024)
Pata de gallo (Spanish) (Rehm 1994)
Pie de gallina (Spanish) (Rehm 1994)
Zacate de agua (Spanish) (Rehm 1994; Rojas-Sandoval and Acevedo-Rodriguez 2014)
Bai (transcribed Chinese 稗) (efloras 2024)

  • Echinochloa crus-galli (barnyard grass) florets

  • Echinochloa crus-galli (barnyard grass) spikelet, floret, and caryopsis

  • Echinochloa crus-galli (barnyard grass) spikelets and partial spikelets showing lemmas

  • Echinochloa crus-galli (barnyard grass) spikelet, floret, and caryopsis

  • Echinochloa crus-galli (barnyard grass) spikelet

  • Echinochloa crus-galli (barnyard grass) spikelet

  • Echinochloa crus-galli (barnyard grass) partial spikelet showing lemma

  • Echinochloa crus-galli (barnyardgrass) fertile floret in palea view.  Note the folded palea tip. Scale in mm.

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Overview

Aperçu

Regulation :

Remarques Réglementation:

    Regulation Notes:

    • Echinochloa crus-galli is a restricted noxious weed seed in the Arkansas, USA, when found as a seed lot contaminant in amounts greater than 300 seed per pound (USDA-AMS 2024).
    • E. crus-galli is listed as a Harmful Organism by Grenada and Panama (USDA-PCIT 2024).
    • E. crus-galli, E. crus-galli var. echinata, and E. crus-galli var. praticola are listed as Harmful Organisms by The Democratic People’s Republic of Korea and The Republic of Korea (USDA-PCIT 2024).

    Distribution :

    Répartition :

    The origin of Echinochloa crus-galli is reported as probably tropical Asia (Rojas-Sandoval and Acevedo-Rodriguez 2014; USDA-ARS 2024). The native range is believed to extend from southern and eastern Europe to Asia, tropical Africa to South Africa, and Madagascar (POWO 2024) or it is native to the western Russia Federation (USDA-ARS 2024). Whatever the origin and native range may be, this species is now widespread throughout warm-temperate, subtropical and tropical regions of the world from 50°N to 40°S latitudes (Rojas-Sandoval and Acevedo-Rodriguez 2014).

    Habitat and Crop Association :

    Habitat et Cultures Associées :

    Echinochloa crus-galli is a widespread weed species found in agricultural fields, moist places, along streambanks, irrigation cannels, ditches, and other disturbed sites (Barkworth et al. 2003; efloras 2024). In agriculture, E. crus-galli is a major weed that can cause significant yield loss in a wide range of crops, such as Glycine max (L.) Merr. (soybean), Gossypium L. spp. (cotton), Medicago sativa L. (lucerne), Sorghum bicolor (L.) Moench (sorghum), Zea mays L. (maize), and many others (Holm et al. 1991; Rojas-Sandoval and Acevedo-Rodriguez 2014). It is especially troublesome in Oryza sativa L. (rice) fields (Holm et al. 1991; Rojas-Sandoval and Acevedo-Rodriguez 2014). This species is also considered an invasive weed in native grasslands, coastal forests, and disturbed areas on most continents (Rojas-Sandoval and Acevedo-Rodriguez 2014).

    Economic Use, cultivation area, and Weed Association :

    Utilisation économique, zone de culture et association de mauvaises herbes :

    Echinochloa crus-galli is used for livestock forage and fodder in many tropical and subtropical regions of the world (Boulos and el-Hadidi 1984; Rojas-Sandoval and Acevedo-Rodriguez 2014).

    Duration of Life Cycle :

    Durée du cycle vital:

    Annual

    Dispersal Unit Type :

    Type d’unité de dispersion :

    Spikelet, floret, caryopsis

    General Information

    RENSEIGNEMENTS GÉNÉRAUX

    Echinochloa is a genus of 40 – 50 species found in tropical to warm-temperate, damp environments. Many of the species tend to intergrade, are highly variable, and can be difficult to distinguish (Barkworth et al. 2003; DiTomaso and Healy 2007; Mohler et al. 2021). E. crus-galli is treated by some authors as a single, highly variable species, while others prefer to subdivide it into an assortment of subspecific taxa based on hair (trichome) length and abundance, awn length, leaf size, inflorescence branching pattern, and spikelet color (Barkworth et al. 2003; eFloras 2024).

    E. crus-galli is believed to be the most widespread species of Echinochloa worldwide (Costea and Tardif 2002).

    Average seed weight of 10 populations of E. crus-galli varied from 1.7 – 2.1 mg (Mohler et al. 2021). A single plant can produce many thousands of seeds that can retain a high viability for up to 3 years when buried in soil, some seeds can survive up to 13 years, although survival rates are quite variable depending on environmental conditions and depth of burial (Mohler et al. 2021). Seeds are dispersed by wind, water, clinging to animals and farm equipment, and as a seed lot contaminant (Rojas-Sandoval and Acevedo-Rodriguez 2014; USDA-ARS 2024).

    Some populations of E. crus-galli in China, Egypt, Italy, Spain, the Philippines, and the United States have been reported as resistant to some common herbicides used in Oryza sativa (rice) and Zea mays (maize) production (Rojas-Sandoval and Acevedo-Rodriguez 2014).

    .

    Echinochloa crus-galli plant in a soybean field (Peter M. Dziuk, Minnesota Wildflowers, MinnesotaWildflowers.info)

    Identification

    Identification

    <
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    • Spikelet

      Size

      • Spikelet length*: 3.9 – 4.7 mm (average 4.2 mm); width*: 1.8 – 2.1 mm (average 2.0 mm).
      • Lower (1st) glume*: 1.5 – 2.0 mm (average 1.8 mm). Lower glume one-third to nearly one-half the length of the spikelet (Costea and Tardif 2002).
      • Upper (2nd) glume length*: 3.9 – 4.7 mm (average 4.2 mm); width*: 1.8 – 2.1 mm (average 2.0 mm). Upper glume equal to length of the spikelet.
      *Note: minimum and maximum based on a random selection of 10 spikelets in normal range of this species using image measurement protocol (ISMA 2020). CDA-S-22874. Measured by D.J. Lionakis Meyer.
      • Measurements from the literature:
        • Spikelet length: 2.8 – 3.4 mm; width 1.6 – 1.8 mm (Costea and Tardif 2002).
        • Spikelet length: 2.5–4.0 mm; width: 1.1–2.3 mm wide (Barkworth et al. 2003; eFloras 2024).
        • Spikelet length: 2.3 – 2.7 mm; width: 1.7 – 1.9 mm (Bojňanský and Fargašová 2007; note incorrectly labeled as ‘floret’ in the text).
        • Spikelet length: 3 – 5 mm; width: 1.1–2.3 mm wide (Walters 2011).

      Shape

      • Spikelet oval, egg-shaped, or teardrop-shaped, dorsoventrally compressed, dorsal side arched and ventral side flat (plano-convex, D-shaped in lateral view).
      • Lower (1st) broadly triangular, the outer corners wrapping around the base of the upper glume.
      • Upper (2nd) glume egg-shaped, tapering to an elongated point at the tip.

      Surface Texture

      • Glumes papery, surface with soft short hairs.
      • Lower (1st) glume 3-nerved (Costea and Tardif 2002).
      • Upper (2nd) glume 5 to 7-nerved (Costea and Tardif 2002), nerves usually with short or long, stiff, spine-like hairs.

      Colour

      • Glumes pale green to light yellowish-brown coloured, sometimes purple tinged.

      Other Features

      • The spikelet consists of a pair of glumes, one sterile floret, and one fertile floret.
      • Disarticulation occurs below the glumes; the spikelet falls as a unit from the plant.
    • Sterile (lower) floret

      Size

      • Sterile lemma length*: 3.2 – 3.8 mm (average 3.6 mm): width*: 1.6 – 2.1 mm (average 3.6 mm). Sterile floret lemma nearly equal to length of spikelet.
      • Sterile palea about equal in length to sterile lemma.
      • Awn of sterile floret lemma, if present, up to 5 cm long (Barkworth et al. 2003).
      *Note: minimum and maximum based on a random selection of 10 sterile lemmas in normal range of this species using image measurement protocol (ISMA 2020). CDA-S-22874. Measured by D.J. Lionakis Meyer.

      Shape

      • Sterile lemma egg-shaped to teardrop shaped, tapering to a folded point.
      • Sterile palea oval, slightly pointed, 2-keeled

      Surface Texture

      • Sterile lemma papery, similar to the upper glume.
      • Sterile lemma 7-nerved, lateral nerves visible along their entire length or only towards the lemma tip (Costea and Tardif 2002).
      • Sterile floret surface covered soft short hairs, nerves usually with short or long, stiff, spine-like hairs (up to 1 mm long; Costea and Tardif 2002).
      • Sterile palea membranous, keels with short fine hairs.

      Colour

      • Sterile lemma pale green to light yellowish-brown coloured, sometimes purple tinged.
      • Sterile palea translucent, whitish.

      Other Features

      • Lower sterile floret (Barkworth et al. 2003).
    • Floret (fertile)

      Size

      • Fertile lemma length*: 2.7 – 3.3 mm (average 3.0 mm); width*: 1.6 – 2.0 mm (average 1.8 mm).
      • Palea nearly equal in length to lemma.
      *Note: minimum and maximum based on a random selection of 10 fertile florets in normal range of this species using image measurement protocol (ISMA 2020). CDA-S-22874. Measured by D.J. Lionakis Meyer.
      • Measurements from the literature:
        Fertile lemma length: 2 – 3.5 mm (Walters 2011).
        Fertile lemma length: 2 – 3 mm (eFloras 2024).

      Shape

      • Fertile floret dorsoventrally compressed, dorsal side arched and ventral side flat (plano-convex, D-shaped in lateral view).
      • Fertile lemma egg-shaped, teardrop-shaped, or oval; margins in-rolled over margin of palea.
      • Fertile lemma tip tapered to a broad point, at maturity the tip appears shriveled or laterally pinched.
      • Fertile palea oval and rounded at tip, palea tip usually folded down like a small flap (MWI 2024).

      Surface Texture

      • Fertile lemma and palea hardened (excluding the tips), glabrous, longitudinally striate.
      • Tip of fertile palea is a small membranous flap of tissue.
      • Tip of fertile lemma thin, papery, and shriveled at maturity.
      • At transition from hard lemma body to shriveled tip is a line of minute hairs.
      • Fertile lemma 5-nerved.

      Colour

      • Fertile lemma and palea glossy, light yellowish-brown to grayish-brown coloured.
      • Nerves of lemma usually light yellowish coloured.
    • Caryopsis

      Size

      • Caryopsis length: 1.2 – 2.4 mm; width: 1.0 – 1.8 mm wide (Barkworth et al. 2003; Walters 2011).

      Shape

      • Caryopsis round to oblong, dorsoventrally compressed.

      Surface Texture

      • Caryopsis surface smooth to wrinkled, longitudinally striate, glabrous.

      Colour

      • Caryopsis body and embryo area light brown, brownish-red or brownish-grey coloured (Costea and Tardif 2002).
      • Hilum reddish brown.

      Other Features

      • Hilum nearly round.
    <
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    <
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    • Embryo

      Size

      • Embryo three-fifths to seven-eights the length of the caryopsis.

      Shape

      • Embryo oval to egg-shaped.

      Endosperm

      • Endosperm solid (Terrell 1977).

      Other Features

      • Embryo in lateral position (Martin 1946).

    Identification Tips

    CONSEILS POUR L’IDENTIFICATION

    Echinochloa crus-galli is in the Paniceae tribe of Poaceae (Barkworth et al. 2003).

    • The natural dispersal unit is a spikelet that has one sterile and one fertile floret.
    • The spikelet is egg-shaped or teardrop-shaped in outline and D-shaped in profile view.
    • The papery glumes are unequal in size.
    • The lower (1st) glume is somewhat triangular, wraps around the upper (2nd) glume, and is less than half as long as the spikelet.
    • The upper (2nd) glume is as long as the spikelet and may have short or long spike-like hairs on the nerves.
    • The lemma of the sterile floret looks similar in size and texture to the upper glume. The sterile lemma may or may not be awned, awn length is variable.
    • The palea of the sterile floret is membranous and translucent.
    • The lemma and palea of the fertile floret are hard, smooth, and shiny.
    • The tip of the fertile lemma has a line of tiny hairs at the transition between the hard and shiny main body and the thin, membranous, and withered tip.

    Additional Botany Information

    AUTRES RENSEIGNEMENTS BOTANIQUES

    Flowers/Inflorescence

    • The inflorescence of Echinochloa crus-galli is often an erect or drooping, pyramid shaped panicle, up to 25 cm long (Costea and Tardif 2002; Barkworth et al. 2003; DiTomaso and Healy 2007).
    • The primary branches are stiff, pointed upwards or slightly spreading, 1.5 – 10 cm long, widely spaced apart near the base of the inflorescence and more crowded towards the top (Barkworth et al. 2003; DiTomaso and Healy 2007).
    • Lower primary branches can have shorter secondary branches (Barkworth et al. 2003; DiTomaso and Healy 2007).
    • Spikelets are irregularly arranged on the branches (Costea and Tardif 2002).
    • Branch axes are glabrous or hairy, hairs up to 5 mm long and may have swollen bases (Barkworth et al. 2003).

    Vegetative Features

    • Echinochloa crus-galli plant morphology is extremely variable (DiTomaso and Healy 2007).
    • E. crus-galli plants have a shallow, fibrous root system (DiTomaso and Healy 2007).
    • Stems up to 2 m tall, erect or lying on the ground (Barkworth et al. 2003).
    • Stem nodes are usually glabrous, lower nodes sometimes hairy (Barkworth et al. 2003).
    • Leaf sheaths glabrous, without ligules and auricles, collar region glabrous, auricle region sometimes long-hairy (DiTomaso and Healy 2007; Rojas-Sandoval and Acevedo-Rodriguez 2014).
    • Leaf blades up to 65 cm long and 3 cm wide, usually glabrous, sometimes sparsely hairy (Barkworth et al. 2003).
    • Sheaths and blades green, sometimes purple tinged near base and along margins (Rojas-Sandoval and Acevedo-Rodriguez 2014).

    Similar Species

    ESPÈCES SEMBLABLES

    Similar species are based on a study of seed morphology of various species, and those with similar dispersal units are identified. The study is limited by physical specimen and literature availability at the time of examination, and possibly impacted by the subjectivity of the authors based on their knowledge and experience. Providing similar species information for seed identification is to make users aware of similarities that could possibly result in misidentification.

    In the genus Echinochloa, most of the important measurable characters overlap somewhat among species (Costea and Tardif 2002). Only the characters considered most useful are described below for separating dispersal units of some similar species from E. crus-galli.

    Echinochloa colona (L.) Link

    [Panicum colonum L. (USDA-ARS 2024); Echinochloa colonum (L.) Link (Boulos and el Hadidi 1984)]
    Jungle rice (AOSA 2024; USDA-NRCS 2024)
    Junglerice (DiTomaso and Healy 2007)
    Jungle ricegrass (USDA-ARS 2024)
    Awnless barnyard grass (Barkworth et al. 2003)

    • The glumes and sterile floret lemma of E. colona are very thin and papery and lack the stiff, spike-like hairs found in E. crus-galli.
    • The sterile floret lemma of E. colona is awnless.
    • The caryopses of E. colona are generally smaller in size (1.2 – 1.6 mm long, 1.0 – 1.3 mm wide) compared to those of E. crus-galli (1.2 – 2.4 mm long, 1 – 1.8 mm wide (Barkworth et al. 2003).
    • The caryopses of E. colona are yellowish or whitish coloured, whereas the caryopses of E. crus-galli are brownish-red or brownish-grey coloured (Costea and Tardif 2002; Barkworth et al. 2003).

    Echinochloa esculenta (A. Braun) H. Schol

    [Panicum esculentum A. Braun); E. crus-galli (L.) P. Beauv. subsp. utilis (Ohwi & Yabuno) T. Koyama; E. utilis Ohwi & Yabuno (Yabuno 1962; USDA-ARS 2024)]
    Japanese barnyard grass (Scholz 1992)
    Japanese millet (Wiersema and Léon 1999; Barkworth et al. 2003; USDA-ARS 2024)

    • The spikelets of E. esculenta are broadly oval, egg-shaped or nearly round in outline and are generally wider (2 – 2.5 mm wide; Barkworth et al. 2003) than those of E. crus-galli (1.1 – 2.3 mm wide; Barkworth et al. 2003).
    • The caryopses in E. esculenta are nearly round in outline, 1.2 – 2.3 mm long and wide, and the embryos are four-fifths to nearly the length of the caryopses, whereas the caryopses in E. crus-galli are usually narrower than they are long, and the embryos are three-fifths to seven-eights the length of the caryopses (Barkworth et al. 2003).
    • E. esculenta is a cultivated species believed to have been originally derived from E. crus-galli about 4000 years ago in Japan (Yabuno 1962; deWet et al. 1983). It is used for cereal grain, fodder, and birdseed (deWet et al. 1983; Barkworth et al. 2003).

    Echinochloa frumentacea Link

    Siberian millet (Barkworth et al. 2003)
    White panic (Barkworth et al. 2003)

    • Mature spikelets of E. frumentacea are usually whitish or light yellow coloured, whereas those of E. crus-galli are pale green to light yellowish-brown coloured, sometimes purple tinged.
    • The glumes and sterile floret lemma of E. frumentacea are very thin and papery and lack the stiff, spike-like hairs found in E. crus-galli.
    • The sterile floret lemma of E. frumentacea is awnless.
    • Although the caryopses of E. frumentacea are similar in size to E. crus-galli, they are yellowish or whitish coloured, whereas the caryopses of E. crus-galli are brownish-red or brownish-grey coloured (Costea and Tardif 2002; Barkworth et al. 2003).
    • E. frumentacea is believed to have been originally derived from E. colona in India, and possibly Africa (Yabuno 1962; deWet et al. 1983; Barkworth et al. 2003). This cultivated species is grown for cereal grain, fodder, beer brewing, and for birdseed (Barkworth et al. 2003).

    Echinochloa muricata (P. Beauv.) Fernald

    American barnyard grass (Barkworth et al. 2003; USDA-ARS 2024)
    Prickly barnyard grass (USDA-ARS 2024)
    Rough barnyard grass (MWI 2024)

    • The embryo of E. muricata is four-fifths to nine-tenths the length of the caryopsis, whereas the embryo of E. crus-galli is only three-fifths to seven-eights the length of the caryopsis (Gould et al. 1972).
    • The body of the fertile lemma in E. muricata is hardened and gradually transitions into a firm, pointed, membranous tip, whereas in E. crus-galli the hardened portion of the fertile lemma is rounded at the apex, sharply differentiated from the thin, withering, membranous tip, and this transition point is delineated by a line of tiny hairs (Gould et al. 1972).
    • The hardened fertile floret palea in E. muricata tapers to a firm point, while in E. crus-galli the hardened portion of the fertile floret palea has a rounded apex that transitions into a soft membranous tissue that usually folds over like a small flap (MWI 2024).

    Echinochloa oryzicola (Vasinger) Vasinger

    [Panicum oryzicola Vasinger; E. crus-galli (L.) P. Beauv. var. oryzicola (Vasinger) Ohwi; E. phyllopogon auct. pl. (USDA-ARS 2024)]
    Late barnyard grass (Barkworth et al. 2003; USDA-ARS 2024)
    Late watergrass (DiTomaso and Healy 2007)

    • In E. oryzicola the lower(1st) glume is about one-half to three-fifths the length of the spikelet, whereas in E. crus-galli, the lower glume is usually not longer than one-half the length of the spikelet.
    • The sterile floret lemma in E. oryzicola is often thickened and somewhat leathery (coriaceous), smooth, shiny, and with a few stiff hairs (rarely papery and with stiff hairs), while the sterile floret lemma of E. crus-galli is papery and has stiff, spine-like hairs along the nerves (Costea and Tardif 2002).
    • The tip of the sterile floret lemma in E. oryzicola tapers to a sharp point, unawned or awn up to 1.5 mm (Barkworth et al. 2003), while in E. crus-galli the awn may be 0 – 50 mm in length.
    • The embryo of E. oryicola can be up to nearly the length of the caryopsis, whereas the embryo in E. crus-galli is shorter (three-fifths to seven-eights the length of the caryopsis) (Barkworth et al. 2003).
    • The stigmas are often persistent on the mature caryopsis of E. oryzicola (Costea and Tardif 2002).
    • E. oryzicola is classified as a harmful organism by The Democratic People’s Republic of Korea and by The Republic of Korea (USDA-PCIT 2024).

    Echinochloa oryzoides (Ard.) Fritsch

    [Panicum oryzoides Ard.; E. crus-galli (L.) P. Beauv. var. oryzoides (Ard.) Lindm.; E. coarctata Kossenko; E. macrocarpa Vasinger; E. phllopogon (Stapf) Stapf ex Kossenko (USDA-ARS 2024)]
    Early barnyard grass (Barkworth et al. 2003)
    Early watergrass (DiTomaso and Healy 2007)
    Hairy millet (USDA-ARS 2024)

    • Nerves on the sterile floret lemma in E. oryzoides are usually visible the entire length of the lemma, while in E. crus-galli the nerves may only be visible towards the tip of the lemma or for the entire length of the lemma (Costea and Tardif 2002).
    • In E. oryzoides the sterile floret lemma is almost always awned, the awns are 5 to 50 mm long, whereas in E. crus-galli the sterile lemma may be unawned or the awn may be up to 50 mm long (Barkworth et al. 2003; DiTomaso and Healy 2007).
    • The caryopses of E. oryzoides are yellowish in colour, while in E. crus-galli the caryopses are brownish-red to brownish-grey in colour (Costea and Tardif 2002).
    • E. oryzoides is classified as a harmful organism by The Democratic People’s Republic of Korea and by The Republic of Korea (USDA-PCIT 2024).

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    Reference(s)

    Référence(s)

    Association of Official Seed Analysts (AOSA). 2024. Rules for Testing Seeds, Vol. 3: Uniform Classification of Weed and Crop Seeds. Association of Official Seed Analysts, Wichita, Kansas.

    Barkworth, M. E. Capels, K. M., Long, S., and Piep, M. B. 2003. Flora of North America, Vo. 25 Magnoliophyta: Commelinidae (in part): Poaceae, part 2. Oxford University Press.

    Bojňanský, V. and Fargašová, A. 2007. Atlas of Seeds and Fruits of Central and East-European Flora: The Carpathian Mountains Region. Springer, Dordrecht, The Netherlands.

    Boulos, L. and el-Hadidi, M. N. 1984. The Weed Flora of Egypt. The American University in Cairo Press.

    Costea, M. and Tardif, F.J. 2002. Taxonomy of the most common weedy European Echinochloa species (Poaceae: Panicoideae) with special emphasis on characters of the lemma and caryopsis. Sida 20(2):525-548.

    Darbyshire, S. J. 2003. Inventory of Canadian Agricultural Weeds Agriculture and Agri-Food Canada. 45.

    de Wet, J. M. J., Prasada Rao, K. E., Mengesha, M. H., and Brink, D. E. 1983. Domestication of sawa millet (Echinochloa colona). Economic Botany 37(3):283-291.

    DiTomaso, J. M. and Healy, E. A.  2007.  Weeds of California and Other Western States.  Vol. 2: Geraniaceae – Zygophyllaceae.  Agriculture and Natural Resources Publication 3488.  University of California, Oakland, CA.

    eFloras. 2024. Flora of China. Echinochloa crusgalli (L.) Beauv. http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=220004563 Accessed September 13, 2024.

    Gould, F. W., Ali, M. A., and Fairbrothers, D. E. 1972. A revision of Echinochloa in the United States. The American Midland Naturalist, 87(1), 36–59. https://doi.org/10.2307/2423880

    Holm, L. G., Plucknett, D. L., Pancho, J. B., Herberger, J. P. 1977. The World’s Worst Weeds: Distribution and Biology. The University Press of Hawaii, Honolulu, HI.

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    Author(s)

    AUTEUR(S)

    Deborah J. Lionakis Meyer

    California Department of Food and Agriculture (retired).