Triticum turgidum subsp. durum
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Overview
Aperçu
Regulation :
Remarques Réglementation:
Regulation Notes:
Distribution :
Répartition :
Habitat and Crop Association :
Habitat et Cultures Associées :
Economic Use, cultivation area, and Weed Association :
Utilisation économique, zone de culture et association de mauvaises herbes :
Durum wheat has a high yield under dry conditions, preferring hot days and cool nights during the growing season. The cultivation area of durum wheat is concentrated in semi-arid regions of the Mediterranean basin, West Asia, North Africa, Southern Europe and North America (Grant et al. 2012).
Triticum turgidum L. subsp. durum (Desf.) van Slageren (durum wheat) accounts for a very small proportion (5-8%) of the total wheat production in the world (Kadkol & Sissons 2015). The European Union has both the largest production and demand of durum wheat, while Canada is the second largest producer and exporter of the crop. In Canada, most of the durum wheat is grown in the province of Saskatchewan and in the United States the majority is produced in North Dakota and Montana (Grant et al. 2012).
Of all wheat species, durum wheat has the hardest caryopsis (grain) (Liu et al. 1996; Wieser et al. 2020) and many varieties have a yellow endosperm making it ideal for pasta (Wiseman 2001; Wieser et al. 2020). The grain’s hard endosperm yields a course yellow semolina flour on milling (Wiseman 2001). The milling of the semolina flour also results in durum wheat flour, which is much finer in consistency (Agrilinkage 2022). Other products made from durum wheat are flat breads, couscous, and bulgur (Liu et al. 1996).
Durum semolina flour generally has a higher protein and gluten content compared to Triticum aestivum subsp. aestivum (common wheat) flour, however due to its low SDSS (sodium dodecyl sulfate sedimentation) content, durum has a much weaker and less elastic gluten resulting in its poor bread making quality (Liu et al. 1996).
Weed surveys from 2009 to 2017 found weed species present in Canadian durum wheat fields with the highest relative abundanceto be: Setaria viridis (L.) P. Beauv. (green foxtail), Avena fatua L. (wild oat), Brassica napus subsp. napus L. (volunteer canola), Thlaspi arvense L. (stinkweed), Fallopia convolvulus (L.) Á. Löve (wild buckwheat), Bassia scoparia (L.) A. J. Scott (kochia), Chenopodium album L. (lamb’s quarters), Cirsium arvense (L.) Scop. (Canada thistle), Salsola tragus L. (Russian thistle), and Sinapis arvensis L. (wild mustard) (Geddes et al. 2020).
Duration of Life Cycle :
Durée du cycle vital:
Annual or winter annual (FNA 1993+)
Dispersal Unit Type :
Type d’unité de dispersion :
Caryopsis
General Information
RENSEIGNEMENTS GÉNÉRAUX
Triticum turgidum subsp. durum is a tetraploid species (AABB) that originated from the Fertile Crescent in the Middle East (Wiseman 2000). Durum wheat is a free threshing wheat (caryopses loosely held within spikelets) that was derived from the hulled (caryopses tightly held within spikelets) Triticum turgidum subsp. dicoccon (Schrank) Thell. (emmer) (Wieser et al. 2020). Hard, translucent, and glass-like (vitreous) caryopses of durum wheat are desired for pasta making, because they provide a high yield of semolina (Kadkol & Sissons 2015). Starchy or non-vitreous kernels produce more flour than semolina resulting in inferior pasta.
.Identification
Identification
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Spikelet
Size
- Size from literature:
• Spikelet length 10 – 15 mm (FNA 1993+)
• Glume length 10 – 12 mm (FOC 1994+)
• Glume length 8 – 12 mm (FNA 1993+)
Shape
- Glumes are prominently keeled with a tooth at one end (FNA 1993+)
Surface Texture
- Glumes are leathery (coriaceous) (FNA 1993+; FOC 1994+)
- Glumes have 5-7 nerves (Feldman and Levy 2023)
Colour
- Glumes yellow, but some varieties are red or blue-black in colour (Feldman and Levy 2023)
Other Features
- Spikelets comprised of 5-7 florets; 2-4 of which may produce caryopses (FNA 1993+)
Triticum turgidum subsp. durum A: glume, B: lemma with awn, C: caryopsis, D: palea, E: palea attached to caryopsis
- Size from literature:
-
Floret
Size
- Lemmas are similar length as glumes (FOC 1994+)
- Lemma length: 10 – 12 mm (FNA 1993+)
- Palea is shorter than lemma (Feldman and Levy 2023)
- Lemma awn (if present) length: up to 23 cm (FNA 1993+)
Shape
- Palea is teardrop-shaped
- Lemma is rounded (not keeled) and is teardrop-shaped
Surface Texture
- Lemma observed to have 10 to 11 nerves, but Feldman and Levy (2023) describe as having 9-15 nerves
- Palea is smooth and membranous
- Palea has fine hairs (palea teeth) along its margins
Colour
- Palea is light yellow and translucent
- Lemma is straw coloured
- Lemma awns are white, red, or black coloured (Feldman and Levy 2023)
Other Features
- Palea does not split when mature (FNA 1993+)
- Lower 2 lemmas have awns (FNA 1993+), but some varieties have no awns (Feldman and Levy 2023)
Triticum turgidum subsp. durum A: glume, B: lemma with awn, C: caryopsis, D: palea, E: palea attached to caryopsis
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Caryopsis
Size
- Caryopsis length: mm; width: mm
*Note: minimum and maximum of caryopses in a normal range of one variety using image measurement (ISMA 2020)
- Measurements from literature
• Caryopsis length: 8-10 mm; width: 3-3.6 mm (Bojňanský and Fargašová 2007)
• Caryopsis length: 6-11 mm; width: 1.8-3.5 mm; depth: 1.8-3.5 mm (ISTA 2021)
• Hilum is almost the full length of the caryopsis (ISTA 2021)
Shape
- Caryopsis is elongated oval with a slight central longitudinal ridge in embryo view, with relatively steep slopes on either side
- Caryopsis side opposite embryo may be relatively flat, concave, or sides along the central groove may be somewhat curved
- Hilum is linear and located in the longitudinal groove (ISTA 2021)
Surface Texture
- Caryopsis has a central groove on the hilum side (opposite of embryo side)
- Caryopsis surface is smooth or with transverse wrinkles
- A pinched ridge surrounds the embryo; sometimes it may not be obvious in embryo view, but when caryopsis is in lateral view, the ridge can normally be seen
- Caryopsis end opposite the embryo generally has a brush of short hairs that may appear to be absent until viewed under magnification
Colour
- Caryopses are light yellow to amber coloured
- Caryopses are sometimes white or red in colour (Feldmen and Levy 2023)
Triticum turgidum subsp. durum (durum wheat) caryopses (Canadian Western Amber Durum – variety A)
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Embryo
Size
- Embryo is approximately ¼ to 1/3 of the caryopsis
Shape
- Embryo is oval to egg-shaped
Endosperm
- Endosperm is generally hard, translucent, and generally yellow but may be white
- Endosperm is flinty, but sometimes mealy (FNA 1993+)
Other Features
- The embryo is located on the caryopsis end opposite the end with hairs and on the side opposite the crease and hilum
- Embryo is in the lateral position
Durum wheat (Triticum turgidum subsp. durum)
Identification Tips
CONSEILS POUR L’IDENTIFICATION
Triticum turgidum subsp. durum generally has an elongated and larger caryopsis with a brush of hairs on the end opposite the embryo that is much reduced or absent compared to Triticum aestivum subsp. aestivum. In embryo view, durum caryopses have a slight central ridge with relatively steep sides on either side and a pinched ridge around all or some of the embryo, which is not present in T. aestivum subsp. aestivum and ×Triticosecale spp..

Durum wheat (Triticum turgidum subsp. durum)









Additional Botany Information
AUTRES RENSEIGNEMENTS BOTANIQUES
Flowers/Inflorescence
- Plants are erect with 3-4 culms and are 60 – 160 cm tall (Feldman and Levy 2023)
- Blades are often glabrous, 7 – 16 mm wide (FNA 1993+), 16 – 25 cm long, with a transparent ligule and auricles that have fine hairs (Feldman and Levy 2023)
Vegetative Features
- Spikes are 4 – 11 cm long (awns not included) and comprised of 20 spikelets on average
Similar Species
ESPÈCES SEMBLABLES
Similar species are based on a study of seed morphology of various species, and those with similar dispersal units are identified. The study is limited by physical specimen and literature availability at the time of examination, and possibly impacted by the subjectivity of the authors based on their knowledge and experience. Providing similar species information for seed identification is to make users aware of similarities that could possibly result in misidentification.
Triticum aestivum L. subsp. aestivum (common wheat)
Triticum aestivum subsp. aestivum caryopses generally have a plump rounded cheek area on either side of the crease which Triticum turgidum subsp. durum often lack. Also, the pinched rim or ridge around the durum wheat embryo is absent in common wheat. The brush of hairs opposite the embryo end are often thicker and more obvious in common wheat. Cross sections of the common wheat caryopsis reveal a white endosperm that may be hard or very mealy, whereas durum wheat is generally yellow in colour and hard (although it can also be white and soft).
×Triticosecale spp. Wittm. ex A. Camus (triticale)
×Triticosecale spp. caryopses have a surface texture generally with more obvious transverse wrinkles than those of T. turgidum subsp. durum. Another feature of ×Triticosecale is that the outer layer of the caryopsis is often flaky and loosely adhered to the internal portion, which is not seen in durum wheat. The embryo of ×Triticosecale spp. generally lacks the pinched ridge around the embryo as in durum wheat and the endosperm is white and not the usual yellow colour of durum. The brush of hairs appears more obvious in ×Triticosecale spp. than durum wheat’s, which may appear to be missing or is comprised of short hairs.
Triticum turgidum L. subsp. dicoccon (Schrank) Thell. (Triticum turgidum Dicoccum Group) (emmer).
T. turgidum subsp. dicoccon is not a free-threshing species and is generally in spikelet form (Wieser et al. 2020). Emmer caryopses are generally long oval shaped with both ends relatively narrow in comparison to durum wheat. The embryo extends further past the margin of the caryopsis in emmer than it does in durum wheat and the end opposite the embryo has a brush of long hairs. The brush of hairs is much more obvious than durum wheats.
Triticum turgidum L. subsp. turanicum (Jakubz.) Á. Löve & D. Löve (kamut; khorasan wheat)
Some authors refer to kamut as Triticum turgidum subsp. durum (FNA 1993+; USDA-ARS 2024), whereas other literature places it as a separate subspecies of Triticum turgidum known as turanicum (Feldman and Levy 2023). Khorasan wheat is often referred to as kamut, which is a common cultivar name (Feldman and Levy 2023). T. turgidum subsp. turanicum caryopses are longer than durum wheat with long straight sides giving it an oblong shape. The profile of kamut’s caryopsis is less humped than durum wheats. The grooved side of the caryopsis often has a slight arch so that when placed lying on this side, a hollow area can be seen.
Triticum monococcum L. subsp. monococcum (einkorn)
T. monococcum subsp. monococcum, like spelt and emmer, is considered a hulled wheat (Longin et al. 2016). T. monococcum subsp. monococcum caryopses are strongly laterally compressed so that the embryos are located on the narrow edges. The embryo of T. monococcum subsp. monococcum noticeably extends further past the margin of the caryopsis than it does in T. turgidum subsp. durum.
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Triticum aestivum subsp. aestivum

Triticosecale spp.

Triticum turgidum subsp. dicoccon

Triticum monococcum subsp. monococcum
Comparison Window
Fenêtre de comparaison
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum (durum wheat) caryopses (Canadian Western Amber Durum – variety A)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum (durum wheat) caryopses (Canadian Western Amber Durum – variety A)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum (durum wheat) caryopses (Canadian Western Amber Durum – variety A)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum (durum wheat) caryopsis (Canadian Western Amber Durum – variety A)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum (durum wheat) caryopsis cross section (Canadian Western Amber Durum – variety A)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum (durum wheat) caryopses (Canadian Eastern Amber Durum)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum (durum wheat) caryopses (Canadian Eastern Amber Durum)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum (durum wheat) caryopses (Canadian Western Amber Durum – variety B)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Durum wheat (Triticum turgidum subsp. durum)
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum A: glume, B: lemma with awn, C: caryopsis, D: palea, E: palea attached to caryopsis
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum spike
MAIN SPECIES
ESPÈCES PRINCIPALES
Triticum turgidum subsp. durum

Triticum turgidum subsp. durum
Poaceae
Triticum turgidum subsp. durum floret in palea view with sterile floret attached
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Common wheat (Triticum aestivum subsp. aestivum)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Common wheat (Triticum aestivum subsp. aestivum)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Common wheat (Triticum aestivum subsp. aestivum)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Common wheat (Triticum aestivum subsp. aestivum)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopsis (Canada Western Soft White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopsis cross-section (Canada Western Soft White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopses (Canada Western Soft White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopses (Canada Western Soft White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopses (Canada Western Soft White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopsis (Canada Western Hard White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopsis cross-section (Canada Western Hard White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopses (Canada Western Hard White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopses (Canada Western Hard White Spring type)
SIMILAR SPECIES
ESPÈCES SEMBLABLES
Triticum aestivum subsp. aestivum

Triticum aestivum subsp. aestivum
Poaceae
Triticum aestivum subsp. aestivum (common wheat) caryopses (Canada Western Hard White Spring type)
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Reference(s)
Référence(s)
Agrilinkage. 2022. https://www.agrilinkage.com/post/semolina-vs-durum-wheat Accessed November 27, 2024.
Bojňanský, V. and Fargašová, A. 2007. Atlas of Seeds and Fruits of Central and East-European Flora: The Carpathian Mountains Region. Springer, Dordrecht, The Netherlands. 1046 pp.
Feldman, M. and Levy, A. 2023. Chapter 10 – Triticum L. In: Feldman, M. and Levy, A. Wheat evolution and domestication. Cham: Springer International Publishing, pp. 365-526. https://link.springer.com/book/10.1007/978-3-031-30175-9#about-authors
Ferrante A., Savin R., Slafer G. A. 2010. Floret development of durum wheat in response to nitrogen availability. Journal of Experimental Botany (61)-15: 4351-4359. https://doi.org/10.1093/jxb/erq236
Flora of China (FOC) 1994+. Triticum turgidum subsp. durum Vol. 22 Page 443 (English edition). http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=250071110 Accessed September 11, 2024.
Flora of North America (FNA) Editorial Committee, eds. 1993+. Flora of North America North of Mexico [Online]. 22+ vols. New York and Oxford. http://beta.floranorthamerica.org Accessed September 11, 2024.
Geddes C. M., Tidemann B. D., Wolf T. & Johnson E. N. 2020. 10. Integrated Weed Management To Minimize Yield Losses in Barilla America Inc. (Pub.), Sustainable Production of Durum Wheat in Canada: 50-57. https://www.albertabarley.com/wp-content/uploads/2020/02/durum_production_manual.pdf
Grant C. A., Fonzo N. D. & Pisante M. 2012. CHAPTER 3 – Agronomy of Durum Wheat Production in M. Sissons, J. Abecassis, B. Marchylo & M. Carcea (Eds.), American Associate of Cereal Chemists International, Durum Wheat (Second Edition): 37-55. https://doi.org/10.1016/B978-1-891127-65-6.50008-8
International Seed Morphology Association (ISMA). 2020. Method for Seed Size Measurement. Version 1.0. ISMA Publication Guide. https://www.idseed.org/authors/details/method_for_seed_size_measurement.html
International Seed Testing Association (ISTA). 2021. Universal List of Species. https://www.seedtest.org/en/services-header/tools/purity-committee/universal-list-species.html
Kadkol G. P. & Sissons M. 2015. Durum Wheat Overview. Reference Module in Food Science (2016). https://doi.org/10.1016/B978-0-08-100596-5.00024-X
Liu C. Y., Kenneth W. & Rathjen A. J. 1996. Improvement of durum wheat pastamaking and breadmaking qualities. Cereal Chemistry (73): 155-166. https://www.cerealsgrains.org/publications/cc/backissues/1996/Documents/73_155.pdf
U.S. Department of Agriculture-Agricultural Research Services (USDA-ARS). 2022. Germplasm Resources Information Network (GRIN), https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomysearch Accessed September 11, 2024.
Wiersema, John, H. and Blanca León. 2016. World Economic Plants. Available from: VitalSource Bookshelf, (2nd Edition). Taylor & Francis.
Wieser, H., Koehler, P. & Scherf, K. A. 2020. Wheat an Exceptional Crop: Botanical Features, Chemistry, Utilization, Nutritional and Health Aspects. https://www.sciencedirect.com/book/9780128217153
Wiseman G. 2001. Chapter 2: Durum Wheat, in R.C. Kill & K. Turnbull (Eds.), Pasta and Semolina Technology: 11-42. https://doi.org/10.1002/9780470999370.ch2